TYPE: Costa Rica: San José: Pérez Zeledón, San Ramón Norte, trail to Cerro Pelón, near the summit of the mountain, 1640 m, 3 May 2000, F. Pupulin, M.L. Spadari, J. Cambronero, V. Juarez-Pérez & K. Granados 2372 (holotype, USJ; detail of type, USJ; illustration of type).